0000001 - Document - Cover Page link:presentationLink link:calculationLink link:definitionLink 9952151 - Statement - Condensed Consolidated Balance Sheets (Unaudited) link:presentationLink link:calculationLink link:definitionLink 9952152 - Statement - Condensed Consolidated Balance Sheets (Unaudited) (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 9952153 - Statement - Condensed Consolidated Statements of Operations (Unaudited) link:presentationLink link:calculationLink link:definitionLink 9952154 - Statement - Condensed Consolidated Statements of Comprehensive Loss (Unaudited) link:presentationLink link:calculationLink link:definitionLink 9952155 - Statement - Condensed Consolidated Statements of Stockholders’ Equity (Unaudited) link:presentationLink link:calculationLink link:definitionLink 9952156 - Statement - Condensed Consolidated Statements of Stockholders’ Equity (Unaudited) (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 9952157 - Statement - Condensed Consolidated Statements of Cash Flows (Unaudited) link:presentationLink link:calculationLink link:definitionLink 9952158 - Disclosure - Nature of the Business link:presentationLink link:calculationLink link:definitionLink 9952159 - Disclosure - Summary of Significant Accounting Policies link:presentationLink link:calculationLink link:definitionLink 9952160 - Disclosure - Cash Equivalents and Marketable Securities link:presentationLink link:calculationLink link:definitionLink 9952161 - Disclosure - Fair Value Measurements link:presentationLink link:calculationLink link:definitionLink 9952162 - Disclosure - Accrued Expenses link:presentationLink link:calculationLink link:definitionLink 9952163 - Disclosure - Commitments and Contingencies link:presentationLink link:calculationLink link:definitionLink 9952164 - Disclosure - Collaboration and License Agreements link:presentationLink link:calculationLink link:definitionLink 9952165 - Disclosure - Common Stock and Preferred Stock link:presentationLink link:calculationLink link:definitionLink 9952166 - Disclosure - Stock-Based Compensation link:presentationLink link:calculationLink link:definitionLink 9952167 - Disclosure - Net Loss per Share link:presentationLink link:calculationLink link:definitionLink 9952168 - Disclosure - Related Party Transactions link:presentationLink link:calculationLink link:definitionLink 9952169 - Disclosure - Segment Reporting link:presentationLink link:calculationLink link:definitionLink 9952170 - Disclosure - Subsequent Events link:presentationLink link:calculationLink link:definitionLink 9955511 - Disclosure - Summary of Significant Accounting Policies (Policies) link:presentationLink link:calculationLink link:definitionLink 9955512 - Disclosure - Cash Equivalents and Marketable Securities (Tables) link:presentationLink link:calculationLink link:definitionLink 9955513 - Disclosure - Fair Value Measurements (Tables) link:presentationLink link:calculationLink link:definitionLink 9955514 - Disclosure - Accrued Expenses (Tables) link:presentationLink link:calculationLink link:definitionLink 9955515 - Disclosure - Common Stock and Preferred Stock (Tables) link:presentationLink link:calculationLink link:definitionLink 9955516 - Disclosure - Stock-Based Compensation (Tables) link:presentationLink link:calculationLink link:definitionLink 9955517 - Disclosure - Net Loss per Share (Tables) link:presentationLink link:calculationLink link:definitionLink 9955518 - Disclosure - Segment Information (Tables) link:presentationLink link:calculationLink link:definitionLink 9955519 - Disclosure - Nature of the Business (Details) link:presentationLink link:calculationLink link:definitionLink 9955520 - Disclosure - Cash Equivalents and Marketable Securities - Investment Profile (Details) link:presentationLink link:calculationLink link:definitionLink 9955520 - Disclosure - Cash Equivalents and Marketable Securities - Investment Profile (Details) link:presentationLink link:calculationLink link:definitionLink 9955521 - Disclosure - Cash Equivalents and Marketable Securities - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955522 - Disclosure - Cash Equivalents and Marketable Securities - Contractual Maturity of Marketable Securities (Details) link:presentationLink link:calculationLink link:definitionLink 9955523 - Disclosure - Fair Value Measurements - Summary of Financial Assets Measured at Fair Value on a Recurring Basis (Details) link:presentationLink link:calculationLink link:definitionLink 9955524 - Disclosure - Accrued Expenses - Summary of Accrued Expenses (Details) link:presentationLink link:calculationLink link:definitionLink 9955525 - Disclosure - Commitments and Contingencies - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955526 - Disclosure - Collaboration and License Agreements - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955527 - Disclosure - Common Stock and Preferred Stock - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955528 - Disclosure - Common Stock and Preferred Stock - Summary of Common Stock Reserved For Future Issuance (Details) link:presentationLink link:calculationLink link:definitionLink 9955529 - Disclosure - Stock-Based Compensation - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955530 - Disclosure - Stock-Based Compensation - Summary of Restricted Stock Activity (Details) link:presentationLink link:calculationLink link:definitionLink 9955531 - Disclosure - Stock-Based Compensation - Summary of Stock Option Activity (Details) link:presentationLink link:calculationLink link:definitionLink 9955532 - Disclosure - Stock-Based Compensation - Schedule of Fair Value of Stock Option Awards on the Grant Date Using the Black-Scholes Option Valuation Model (Details) link:presentationLink link:calculationLink link:definitionLink 9955533 - Disclosure - Stock-Based Compensation - Summary of Stock-Based Compensation Expense (Details) link:presentationLink link:calculationLink link:definitionLink 9955534 - Disclosure - Net Loss per Share - Summary of antidilutive securities excluded from computation of earnings per share (Details) link:presentationLink link:calculationLink link:definitionLink 9955535 - Disclosure - Segment Reporting - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955536 - Disclosure - Segment Information - Schedule of Segment Reporting Information, by Segment (Details) link:presentationLink link:calculationLink link:definitionLink